Eight further Achaemenid rulers followed Darius. His two immediate successors, Xerxes (ruled 486-465 BC) and Artaxerxes I (465-424 BC), carried on his building programme. Hostile relations with Greece led to the Graeco-Persian Wars, although territorial acquisitions continued to be made in other areas. The Achaemenid Empire fell to Alexander the Great and his Macedonian army in 330 BC.